Fraxinus Nigra
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''Fraxinus nigra'', the black ash, is a species of ash native to much of eastern Canada and the northeastern United States, from western
Newfoundland Newfoundland and Labrador (; french: Terre-Neuve-et-Labrador; frequently abbreviated as NL) is the easternmost province of Canada, in the country's Atlantic region. The province comprises the island of Newfoundland and the continental region ...
west to southeastern
Manitoba Manitoba ( ) is a Provinces and territories of Canada, province of Canada at the Centre of Canada, longitudinal centre of the country. It is Canada's Population of Canada by province and territory, fifth-most populous province, with a population o ...
, and south to
Illinois Illinois ( ) is a U.S. state, state in the Midwestern United States, Midwestern United States. Its largest metropolitan areas include the Chicago metropolitan area, and the Metro East section, of Greater St. Louis. Other smaller metropolita ...
and northern
Virginia Virginia, officially the Commonwealth of Virginia, is a state in the Mid-Atlantic and Southeastern regions of the United States, between the Atlantic Coast and the Appalachian Mountains. The geography and climate of the Commonwealth ar ...
. Formerly abundant, as of 2014 the species is threatened with near total extirpation throughout its range, as a result of infestation by a parasitic insect known as the
emerald ash borer The emerald ash borer (''Agrilus planipennis''), also known by the acronym EAB, is a green buprestid or jewel beetle native to north-eastern Asia that feeds on ash species. Females lay eggs in bark crevices on ash trees, and larvae feed undern ...
.


Description

Black ash is a medium-sized
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
reaching (exceptionally ) tall with a trunk up to diameter, or exceptionally to . The bark is grey, thick and corky even on young trees, becoming scaly and fissured with age. The winter buds are dark brown to blackish, with a velvety texture. The
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
are opposite, pinnately compound, with 7–13 (most often 9) leaflets; each leaf is long, the leaflets long and broad, with a finely toothed margin. The leaflets are
sessile Sessility, or sessile, may refer to: * Sessility (motility), organisms which are not able to move about * Sessility (botany), flowers or leaves that grow directly from the stem or peduncle of a plant * Sessility (medicine), tumors and polyps that ...
, directly attached to the
rachis In biology, a rachis (from the grc, ῥάχις [], "backbone, spine") is a main axis or "shaft". In zoology and microbiology In vertebrates, ''rachis'' can refer to the series of articulated vertebrae, which encase the spinal cord. In this c ...
without a petiolule. The
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
s are produced in spring shortly before the new leaves, in loose
panicle A panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit) be pedicellate (having a single stem per flower). The branches of a panicle are of ...
s; they are inconspicuous with no petals, and are wind-pollinated. The
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
is a
samara Samara ( rus, Сама́ра, p=sɐˈmarə), known from 1935 to 1991 as Kuybyshev (; ), is the largest city and administrative centre of Samara Oblast. The city is located at the confluence of the Volga and the Samara (Volga), Samara rivers, with ...
long comprising a single
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
long with an elongated apical wing long and broad.New Brunswick tree and shrub
''Fraxinus nigra''
/ref>Virtual Herbarium of the Chicago Region
''Fraxinus nigra''


Ecology and conservation status

Black ash commonly occurs in
swamp A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
s, often with the closely related
green ash ''Fraxinus pennsylvanica'', the green ash or red ash, is a species of ash native to eastern and central North America, from Nova Scotia west to southeastern Alberta and eastern Colorado, south to northern Florida, and southwest to Oklahoma and e ...
. Its fall foliage is yellow. Black ash is one of the first trees to lose its leaves in the fall. It is very closely related to Manchurian ash, and will easily hybridize with it. Some consider the two to be geographic isolates of each other. The species was considered abundant and its survival of little concern prior to the invasion of the
emerald ash borer The emerald ash borer (''Agrilus planipennis''), also known by the acronym EAB, is a green buprestid or jewel beetle native to north-eastern Asia that feeds on ash species. Females lay eggs in bark crevices on ash trees, and larvae feed undern ...
, first detected in North America in 2002. However, since that time this invasive insect has spread throughout most of the tree's range, and within a few years black ash is expected to be all but extirpated; a similar fate awaits green ash. In 2014, a U.S. Forest Service agent estimated that "ninety-nine percent of the ashes in North America are probably going to die."
Blue ash ''Fraxinus quadrangulata'', the blue ash, is a species of ash native primarily to the Midwestern United States from Oklahoma to Michigan, as well as the Bluegrass region of Kentucky and the Nashville Basin region of Tennessee. Isolated populati ...
and white ash are only slightly less affected.


Direct usefulness for humans

This wood is used by Native Americans of the North East for making baskets and other devices. The
Shakers The United Society of Believers in Christ's Second Appearing, more commonly known as the Shakers, are a Millenarianism, millenarian Restorationism, restorationist Christianity, Christian sect founded in England and then organized in the Unit ...
also made extensive use of the black ash for creating baskets. Also called basket ash, brown ash, swamp Ash, hoop ash, and water ash. It is also a popular wood for making electric guitars and basses, due to its good resonant qualities.


Creating basket strips

Black ash is unique among all trees in North America in that it does not have fibers connecting the growth rings to each other. This is a useful property for basket makers. By pounding on the wood with a mallet, the weaker spring wood layer is crushed, allowing the tougher and darker summer wood layer to be peeled off in long strips. The long strips are trimmed, cleaned, and used in
basket weaving Basket weaving (also basketry or basket making) is the process of weaving or sewing pliable materials into three-dimensional artifacts, such as baskets, mats, mesh bags or even furniture. Craftspeople and artists specialized in making baskets ...
.
Indigenous peoples of the Northeastern Woodlands Indigenous peoples of the Northeastern Woodlands include Native American tribes and First Nation bands residing in or originating from a cultural area encompassing the northeastern and Midwest United States and southeastern Canada. It is part ...
also make bark baskets from black ash, traditionally used for berry-gathering.


Usefulness to wildlife

North American native ash tree species are used by North American frogs as a critical food source, as the leaves that fall from the trees are particularly suitable for tadpoles to feed upon in ponds (both temporary and permanent), large puddles, and other water sources. Species such as red maple, which are taking the place of ash, due to the ash borer, are much less suitable for the frogs as a food source—resulting in poor frog survival rates and small frog sizes. It is the lack of tannins in the American ash variety that makes them good for the frogs as a food source and also not resistant to the ash borer. Varieties of ash from outside North America typically have much higher tannin levels and resist the borer. Maples and various non-native invasive trees, trees that are taking the place of American ash species in the North American ecosystem, typically have much higher leaf
tannin Tannins (or tannoids) are a class of astringent, polyphenolic biomolecules that bind to and precipitate proteins and various other organic compounds including amino acids and alkaloids. The term ''tannin'' (from Anglo-Norman ''tanner'', ...
levels. Ash species native to North America also provide important habitat and food for various other creatures that are native to North America, such as the long-horned beetle, avian species, and mammalian species. Black ash is a food plant for the
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...
e of several species of
Lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...
; see
List of Lepidoptera that feed on ashes Ashes (''Fraxinus'' species) are used as food plants by the larvae (caterpillars) of a number of Lepidoptera species, including the following. (Unless otherwise stated, records are from the Nearctic region.) 425px, ''Fraxinus americana'' Adelida ...
.


References

{{Taxonbar, from=Q1790101 nigra Trees of the North-Central United States Trees of the Northeastern United States Trees of the Southeastern United States Trees of Western Canada Trees of Eastern Canada Trees of humid continental climate Least concern plants Trees of Ontario Trees of the Great Lakes region (North America)